Extant and fossil species of Karumballichirus (Decapoda: Axiidea: Callichiridae)

Two extant species of Indo-West Pacific ghost shrimps of the genus Karumballichirus Poore, Dworschak, Robles, Mantelatto and Felder, 2019 that had been previously synonymized, are distinguished on the basis of molecular and morphological criteria. Karumballichirus karumba (Poore and Griffin, 1979) from northern Australia, Papua New Guinea, Singapore, and Thailand differs from Karumballichirus kempi (Sakai, 1999) from India, Indonesia and Taiwan in the shape of the adult cheliped, telson, and uropods, and in reaching little over half the length of K. kempi. Ten fossil species of Karumballichirus, six in new combinations, dating from the Paleocene (Danian) to Holocene are listed: Karumballichirus birmanica (Noetling, 1901) n. comb., Miocene; Karumballichirus dijki (Martin, 1883) n. comb., Miocene; Karumballichirus lakhraensis (Hyžný and Charbonnier in Hyžný, Charbonnier, Merle, Lashari, Bartolini and Métais, 2016), early Eocene (Ypresian); Karumballichirus khadroensis (Hyžný and Charbonnier in Hyžný, Charbonnier, Merle, Lashari, Bartolini and Métais, 2016), Paleocene (Danian); Karumballichirus pseudoniloticus (Lőrenthey in Lőrenthey and Beurlen, 1929) n. comb., middle Eocene; Karumballichirus pustulatus (Withers, 1926) n. comb., ?Eocene-Oligocene; Karumballichirus trechmanni (Withers, 1924) n. comb., Eocene; Karumballichirus tuberculatus (Lőrenthey in Lőrenthey and Beurlen, 1929), middle Eocene; Karumballichirus vidali (Vía Boada, 1959) n. comb., early Eocene, including the subfossil Karumballichirus maximus (A. Milne-Edwards, 1870) which is not a senior synonym of any modern species.