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In a previous three gene phylogeny study (nuITS rDNA, nuLSU rDNA and RPB 1 region A–D) a well supported clade, most closely related to Verrucaria rupestris and also close to Sporodictyon and Atla , was found. Here this clade is identified as Henrica . A phylogeny of these genera based on the nuITS rDNA is supplied, and also a key to crustose species in Verrucariaceae with large, brown muriform spores. Based on material from northern Europe, Henrica is emended to include H. theleodes and H. melaspora , both new combinations in the previously monotypic genus. Verrucaria theleodes and V. scotinospora are lectotypified.
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Savić et al. (2008) studied this question.
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