David G. Lloyd, Jocelyn M. A. Yates, Intrasexual Selection and the Segregation of Pollen and Stigmas in Hermaphrodite Plants, Exemplified by Wahlenbergia albomarginata (Campanulaceae), Evolution, Vol. 36, No. 5 (Sep., 1982), pp. 903-913
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Lloyd et al. (1982) studied this question.
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