The genusInoderma(Ach.) Gray is lectotypified withInoderma byssaceumand resurrected for a small group of species inArthoniaceaewith elevated, white pruinose pycnidia, immersed to adnate white pruinose apothecia and a weakly gelatinized hymenium.Inoderma nipponicumis described from Japan,I. afromontanumfrom Uganda and the EuropeanLecanactis subabietinais transferred to the genus.Sporodophoronis described for a small group of species inArthoniaceaerelated toInodermabut with a unique type of sporodochia instead of pycnidia. Sterile specimens of this new genus resemble species ofTylophoronbut differ in the rounded angular to elliptical to short cylindrical, 0–2-septate sporodochial conidia with unevenly thickened walls which are formed apically in zigzag-shaped and occasionally branched chains.Sporodophoronfurther differs in thallus chemistry and is genetically distinct.Tylophoron americanumandSchismatomma cretaceumare transferred toSporodophoron, and the new speciesS. gossypinumfrom Japan andS. primorskiensefrom eastern Russia are described. The genusGlomerulophoronis described for a single species from Mauritius,G. mauritiae, differing fromSporodophoronin the tightly coiled chains of sporodochial conidia and being genetically distinct. A phylogenetic tree showing the position ofInoderma,SporodophoronandGlomerulophoroninArthoniaceaeis presented. A key to all species ofArthoniaceaewith sporodochia or elevated white pruinose pycnidia is presented.Arthothelium spectabile, the type of the large heterogeneous genusArthothelium, is confirmed for the cryptothecioid subclade inArthoniaceae.
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Frisch et al. (2015) studied this question.
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