A. L. Shafer, J. B. Katz, K. A. Eernisse, Development and Validation of a Competitive Enzyme-Linked Immunosorbent Assay for Detection of Type A Influenza Antibodies in Avian Sera, Avian Diseases, Vol. 42, No. 1 (Jan. - Mar., 1998), pp. 28-34
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Shafer et al. (1998) studied this question.