S. C. L. Fogden, John Proctor, Notes on the Feeding of Land Leeches (Haemadipsa zeylanica Moore and H. picta Moore) in Gunung Mulu National Park, Sarawak, Biotropica, Vol. 17, No. 2 (Jun., 1985), pp. 172-174
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Fogden et al. (1985) studied this question.