Mones S. Abu-Asab, Philip D. Cantino, Systematic Implications of Pollen Morphology in Subfamilies Lamioideae and Pogostemonoideae (Labiatae), Annals of the Missouri Botanical Garden, Vol. 81, No. 4 (1994), pp. 653-686
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Abu‐Asab et al. (1994) studied this question.
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