In crested penguins, Eudyptes spp., the first-laid (A) egg is smaller than the second-laid (B) egg and although viable rarely survives to hatching. I studied survival and growth to fledging of A- and B-chicks at experimentally-manipulated nests, and individual variation in relative investment by adult females in A- and B-eggs, in macaroni penguins, E. chrysolophus
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Tony D. Williams (1990) studied this question.
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