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Saitoh, Kenji, Sado, Tetsuya, Doosey, Michael H., Bart Jr, Henry L., Inoue, Jun G., Nishida, Mutsumi, Mayden, Richard L., Miya, Masaki (2011): Evidence from mitochondrial genomics supports the lower Mesozoic of South Asia as the time and place of basal divergence of cypriniform fishes (Actinopterygii: Ostariophysi). Zoological Journal of the Linnean Society 161 (3): 633-662, DOI: 10.1111/j.1096-3642.2010.00651.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2010.00651.x
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Saitoh et al. (2011) studied this question.
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