Karen T. Coschigano, Rosana Melo-Oliveira, Jackie Lim, Gloria M. Coruzzi, Arabidopsis gls Mutants and Distinct Fd-GOGAT Genes: Implications for Photorespiration and Primary Nitrogen Assimilation, The Plant Cell, Vol. 10, No. 5 (May, 1998), pp. 741-752
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Coschigano et al. (1998) studied this question.