(RMR) HAWAII INSTITUTE OF MARINE BIOLOGY, PO Box 1346, KANEOHE, HAWAII 96744; (TFH): NATIONAL INSTITUTE FOR BASIC BIOLOGY, 38 NISHIGONAKA, MYODAIJICHO, OKAZAKI, 444 JAPAN; (IS): DEPARTMENT OF ZOOLOGY, NATIONAL UNIVERSITY OF SINGAPORE, KENTRIDGE-0511, SINGAPORE. PRESENT ADDRESS (RMR): NATIONAL FISHERY RESEARCH AND DEVELOPMENT LABORATORY, R.D. #4, Box 63, WELLSBORO, PENNSYLVANIA 16901. Accepted 14 June 1989.
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Greenfield et al. (1990) studied this question.