Monique Halloy, Richard Etheridge, Gordon M. Burghardt, To Bury in Sand: Phylogenetic Relationships among Lizard Species of the Boulengeri Group, Liolaemus (Reptilia: Squamata: Tropiduridae), Based on Behavioral Characters, Herpetological Monographs, Vol. 12 (1998), pp. 1-37
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Halloy et al. (1998) studied this question.
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