Eliana A. Simabukuro, Luciano M. Esteves, Gil M. Felippe, Fern Spore Morphology and Spore Rain of a Preserved Cerrado Region in Southeast Brazil (Reserva Biológica e Estação Experimental de Moji Guaçu, São Paulo), American Fern Journal, Vol. 88, No. 3 (Jul. - Sep., 1998), pp. 114-137
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Simabukuro et al. (1998) studied this question.
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