Jorge V. Crisci, Elizabeth A. Zimmer, Peter C. Hoch, George B. Johnson, Christy Mudd, N. S. Pan, Phylogenetic Implications of Ribosomal DNA Restriction Site Variation in the Plant Family Onagraceae, Annals of the Missouri Botanical Garden, Vol. 77, No. 3 (1990), pp. 523-538
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Crisci et al. (1990) studied this question.
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