The heart of Leiopotherapon unicolor comprises 0.135±0.005% of total body weight compared to the average teleost heart size of 0.081±0.002% body weight. The respiratory surface area of 569 mm2 g−1 is also much larger than that found in most freshwater fish. The exponent b in the equation Gill Area =a. body wtb was evaluated at 1.04, whereas the theoretical value for b is only 0.75. It is therefore evident that L. unicolor has an excessively large gill area and a large heart, which facilitate oxygen uptake in hypoxic environments.
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Peter C. Gehrke (1987) studied this question.
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