Muthugapatti K. Kandasamy, Mary K. Thorsness, Sabine J. Rundle, Michael L. Goldberg, June B. Nasrallah, Mikhail E. Nasrallah, Ablation of Papillar Cell Function in Brassica Flowers Results in the Loss of Stigma Receptivity to Pollination, The Plant Cell, Vol. 5, No. 3 (Mar., 1993), pp. 263-275
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Kandasamy et al. (1993) studied this question.