Michael J. Rabe, Thomas E. Morrell, Heather Green, James C. deVos, Jr., C. Richard Miller, Characteristics of Ponderosa Pine Snag Roosts Used by Reproductive Bats in Northern Arizona, The Journal of Wildlife Management, Vol. 62, No. 2 (Apr., 1998), pp. 612-621
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Rabe et al. (1998) studied this question.