Julie E. Gray, Bruce A. McClure, , Marilyn A. Anderson, Adrienne E. Clarke, Action of the Style Product of the Self-Incompatibility Gene of Nicotiana alata (S-RNase) on in Vitro-Grown Pollen Tubes, The Plant Cell, Vol. 3, No. 3 (Mar., 1991), pp. 271-283
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Gray et al. (1991) studied this question.