Joerg Kinne, Mansoor Ali, Ulrich Wernery, J. P. Dubey, Clinical Large Intestinal Coccidiosis in Camels (Camelus dromedarius) in the United Arab Emirates: Description of Lesions, Endogenous Stages, and Redescription of Isospora orlovi, Tsygankov, 1950 Oocysts, The Journal of Parasitology, Vol. 88, No. 3 (Jun., 2002), pp. 548-552
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Kinne et al. (2002) studied this question.