Charles J. Gagen, Richard W. Standage, Joseph N. Stoeckel, Ouachita Madtom (Noturus lachneri) Metapopulation Dynamics in Intermittent Ouachita Mountain Streams, Copeia, Vol. 1998, No. 4 (Dec. 30, 1998), pp. 874-882
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Gagen et al. (1998) studied this question.
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