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Keith S. Delaplane, Arnold M. Saxton, Jeffrey P. La Fage, Foraging Phenology of the Formosan Subterranean Termite (Isoptera: Rhinotermitidae) in Louisiana, The American Midland Naturalist, Vol. 125, No. 2 (Apr., 1991), pp. 222-230
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Delaplane et al. (1991) studied this question.
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