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D. U. U. Okali, J. B. Hall, G. W. Lawson, Root Distribution under a Thicket Clump on the Accra Plains, Ghana: Its Relevance to Clump Localization and Water Relations, Journal of Ecology, Vol. 61, No. 2 (Jul., 1973), pp. 439-454
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Okali et al. (1973) studied this question.
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