J. W. Arntzen, Some Hypotheses on Postglacial Migrations of the Fire-Bellied Toad, Bombina bombina (Linnaeus) and the Yellow-Bellied Toad, Bombina variegata (Linnaeus), Journal of Biogeography, Vol. 5, No. 4 (Dec., 1978), pp. 339-345
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Jan W. Arntzen (1978) studied this question.
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