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John F. Howell, Gonyaulax monilata, sp. nov., the Causative Dinoflagellate of a Red Tide on the East Coast of Florida in August-September, 1951, Transactions of the American Microscopical Society, Vol. 72, No. 2 (Apr., 1953), pp. 153-156
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John F. Howell (1953) studied this question.
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