John C. Clamp, A New Species of Lagenophrys (Ciliophora: Peritricha: Lagenophryidae) Ectocommensal on North American Species of Gammarus (Crustacea: Amphipoda), Transactions of the American Microscopical Society, Vol. 109, No. 2 (Apr., 1990), pp. 121-128
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John C. Clamp (1990) studied this question.
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