John A. Elix, Progress in the Generic Delimitation of Parmelia Sensu Lato Lichens (Ascomycotina: Parmeliaceae) and a Synoptic Key to the Parmeliaceae, The Bryologist, Vol. 96, No. 3 (Autumn, 1993), pp. 359-383
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John Alan Elix (1993) studied this question.
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