A generalized Donnan theory (assuming ions to be adsorbed at sites) is developed to describe the titration curve of an insoluble protein which possesses unequal numbers of carboxyl and amino groups. The equations derived apply to combination with acid or alkali and include a treatment of the isoelectric point. From the constancy of the isoelectric pH of keratin it is concluded that K+ and Cl− do not combine with keratin.
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Luis P. Peters (1960) studied this question.
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