Evergreen or deciduous trees or shrubs, usually with stellate (or lepidote) pubescence.Leaves simple, alternate, exstipulate, with entire, serrate, dentate, or occasionally lobulate margins, the venation pinnate.Flowers perfect, actinomorphic ; inflorescences often reduced, the flowers usually arranged in simple or sometimes branched, terminal and/or axillary racemes, corymbs, or panicles, occasionally solitary or in few-flowered fascicles.Calyx often articulated at the base, synsepalous, tubular, subentire or 4-or 5-toothed or -lobed; calyx tube entirely free to completely adnate to the ovary.Corolla sympetalous, 4-or 5-lobed (rarely more), often united into a tube only at the base; insertion of the corolla hypogynous (ovary superior) to epigynous (ovary inferior), the lobes valvate or imbricate in bud.Stamens usually twice the number of the corolla lobes, sometimes more, epipetalous, inserted in 1 whorl on the base of the corolla (or rarely at the base of the ovary), the filaments usually connate into a tube below, free above; anthers 4-locular.Gynoecium syncarpous, treatments of cultivated ligneous plants, the preparation of which is a project of the Arnold Arboretum of Harvard University and the purpose of which is to in the cooler temperate regions of North America.It is hoped that these treatments will eventually form the basis of a new manual of cultivated woody plants.The first paper in this series was published in the Journal of the Arnold Arboretum 56: 1-19.1975.Reference should be made to the introductory paragraphs of that paper this project, I am especially indebted to Drs.R. A. Howard,' B. G. Schubert, P. F. Stevens, C. E. Wood, Jr., and R. E. Weaver, Jr.I should also like to thank Dr. L. M. Perry for her invaluable help with numerous problems, Mrs. I. H. Burch for her interest in and preliminary studies of Halesia and Pterostyrax in the living collections of the Arnold Arboretum, and Miss K. Clagett for her editorial assistance.The illustrations are the careful work of Robin S. Lefberg and were prepared, when possible, from living or alcohol-preserved specimens collected in the Arnold Arboretum.These collections were made by R. E. Weaver, Jr., M. Gilmore, L H. Burch, and myself.Additional alcohol-preserved specimens collected by C. E. W'ood, Jr., and R. B. Channell from wild populations and from olants in the Uvinir collections of the Henry Foundation for ized.These materials w special tnanks and gratitude are also extended to an anonymous donor, whose
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Stephen A Spongberg (1976) studied this question.