W. Ronald Heyer, A. Stanley Rand, Foam Nest Construction in the Leptodactylid Frogs Leptodactylus pentadactylus and Physalaemus pustulosus (Amphibia, Anura, Leptodactylidae), Journal of Herpetology, Vol. 11, No. 2 (Apr. 25, 1977), pp. 225-228
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Heyer et al. (1977) studied this question.
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