T. Scott Byington, Kurt W. Gottschalk, James B. McGraw, Within-Population Variation in Response of Red Oak Seedlings to Herbivory by Gypsy Moth Larvae, The American Midland Naturalist, Vol. 132, No. 2 (Oct., 1994), pp. 328-339
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Byington et al. (1994) studied this question.
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