Cathy A. Paris, Florence S. Wagner, Warren H. Wagner, Jr., Cryptic Species, Species Delimitation, and Taxonomic Practice in the Homosporous Ferns, American Fern Journal, Vol. 79, No. 2, Species Concepts in Pteridophytes (Apr. - Jun., 1989), pp. 46-54
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Paris et al. (1989) studied this question.
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