G.H. Harper, A. Marchant, D.G. Boddington, The Ecology of the Hen Flea Ceratophyllus gallinae and the Moorhen Flea Dasypsyllus gallinulae in Nestboxes, Journal of Animal Ecology, Vol. 61, No. 2 (Jun., 1992), pp. 317-327
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Harper et al. (1992) studied this question.
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