C. R. Solis, J. E. Strassmann, Presence of Brood Affects Caste Differentiation in the Social Wasp, Polistes exclamans Viereck (Hymenoptera: Vespidae), Functional Ecology, Vol. 4, No. 4 (1990), pp. 531-541
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Solís et al. (1990) studied this question.
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