Sumie Ishiguro, Akiko Kawai-Oda, Junichi Ueda, Ikuo Nishida, Kiyotaka Okada, The DEFECTIVE IN ANTHER DEHISCENCE1 Gene Encodes a Novel Phospholipase A1 Catalyzing the Initial Step of Jasmonic Acid Biosynthesis, Which Synchronizes Pollen Maturation, Anther Dehiscence, and Flower Opening in Arabidopsis, The Plant Cell, Vol. 13, No. 10 (Oct., 2001), pp. 2191-2209
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Ishiguro et al. (2001) studied this question.