Despite the abundance of dinosaur footprints from the Middle Jurassic Cleveland Basin of Yorkshire, UK, examples of footprints indicating two or more different species inhabiting a particular site and environment at the same time are much rarer. The examples described here are of two contrasting dinosaur types, with a single footprint of a large herbivore (ornithopod) and several prints of smaller carnivores (theropod) that were closely associated in the same area. Evidence suggests these tracks were probably made contemporaneously. The reason for their association and suggested behaviour is tentatively suggested, including a potential predator–prey scenario.
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Romano et al. (2026) studied this question.
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