A. Milne, R. Laughlin, R. E. Coggins, The 1955 and 1959 Population Crashes in the Leatherjacket, Tipula paludosa Meigen, in Northumberland, Journal of Animal Ecology, Vol. 34, No. 3 (Oct., 1965), pp. 529-544
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Milne et al. (1965) studied this question.
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