Dave F. Zirkle, Peter S. Dawson, Batia Lavie, An Experimental Analysis of the Genetic Relationships among Life-History Traits and Emigration Behavior in Tribolium Castaneum, Oikos, Vol. 53, No. 3 (Nov., 1988), pp. 391-397
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Zirkle et al. (1988) studied this question.
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