John L. Innes, The Size-Frequency Distributions of the Lichens Sporastatia testudinea and Rhizocarpon alpicola Through Time at Storbreen, South-West Norway, Journal of Biogeography, Vol. 13, No. 4 (Jul., 1986), pp. 283-291
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John L. Innes (1986) studied this question.
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