Masafumi Matsui, Taeko Utsunomiya, Mating Call Characteristics of the Frogs of the Subgenus Babina with Reference to Their Relationship with Rana adenopleura, Journal of Herpetology, Vol. 17, No. 1 (Mar., 1983), pp. 32-37
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Matsui et al. (1983) studied this question.