These observations were made during a fifteen months' tour in Nigeria. From May 1937 till February 1938 the writer was stationed at Kafanchan, 9° 40 N., 8° 20 E., in Plateau Province, with a touring area which included the Jemaa and Southern Divisions of Plateau Province and the Nasarawa Division of BenueProvince; thereafter from March till August he was engaged on sleeping sickness work, which involved extensive travelling in Plateau, Benue, and Niger Provinces.
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William Serle (1939) studied this question.