Nan Zhou, Tina L. Tootle, Frank Tsui, Daniel F. Klessig, Jane Glazebrook, PAD4 Functions Upstream from Salicylic Acid to Control Defense Responses in Arabidopsis, The Plant Cell, Vol. 10, No. 6 (Jun., 1998), pp. 1021-1030
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Tootle et al. (1998) studied this question.