G. F. Midgley, W. D. Stock, J. M. Juritz, Effects of Elevated CO2 on Cape Fynbos Species Adapted to Soils of Different Nutrient Status: Nutrient and CO2-Responsiveness, Journal of Biogeography, Vol. 22, No. 2/3, Terrestrial Ecosystem Interactions with Global Change, Volume 1 (Mar. - May, 1995), pp. 185-191
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Midgley et al. (1995) studied this question.
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