Johan B. Steen, Harald Steen, Nils Chr. Stenseth, Svein Myrberget, , Microtine Density and Weather as Predictors of Chick Production in Willow Ptarmigan, Lagopus l. lagopus, Oikos, Vol. 51, No. 3 (Mar., 1988), pp. 367-373
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Steen et al. (1988) studied this question.
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