Intergeneric hybridizations were performed between four Hordeum and five Asiatic Elymus species, and the embryos were rescued. Meiotic pairing was studied in the parental species and hybrids. The study revealed: i) the allopolyploid nature of the five Asiatic Elymus species; ii) that E. semicostatus (2n=4x=28), E. parviglume (2n=4x=28), and E. pendulinus (2n=4x=28) had no chromosome homology with the Hordeum species; iii) that E. tsukushiensis var. transiens contained a modified H genome from H. hogdanii. thus, its genome symbol should be SSHHYY.
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Lu et al. (2008) studied this question.
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