Study region Alpine headwaters of the Kaidu River (central Tianshan) and Keriya River (northern Kunlun), Northwest China. Study focus We evaluated seasonal performance and bidirectional zero-shot transfer of seven data-driven models and an ensemble with validation-optimized weights for current-month runoff simulation. Models used concurrent and antecedent hydrometeorological and land surface inputs without forecast lead time. Analyses covered seasonal diagnostics, a monthly climatology benchmark, lookback tests (3, 6, 9 months) for four deep learning models, and controlled comparisons of mean squared error (MSE) with hydrology-aware loss. Transfer used fixed source-basin configurations. New hydrological insights for the region Input windows varied by basin and architecture: all four Keriya networks favored 3 months versus 6–9 in Kaidu. These preferences reflect input sensitivity, not measured storage timescales. In Keriya, monthly climatology nearly matched the ensemble in Nash–Sutcliffe efficiency (NSE: 0.791 versus 0.804). Negative NSE in small, low-variance snowmelt and baseflow subsets coincided with modest absolute errors. Hydrology-aware loss improved NSE for Kaidu LSTM and DLinear but reduced it in six other comparisons. Transfer exposed another limitation: ensemble NSE was 0.722 from Keriya to Kaidu and 0.357 in reverse. During transfer to Kaidu, XGBoost exceeded the ensemble (NSE = 0.750), whereas TCN retained high correlation ( r = 0.930) but exaggerated simulated variability (ratio = 2.06; NSE = −0.417).
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