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It has long been a mystery how Fisher (1930, 1941, 1958) derived his famous ‘fundamental theorem of Natural Selection ’ and exactly what he meant by it. He stated the theorem in these words (1930, p. 35; 1958, p. 37): ‘The rate of increase inJitness of any organism at any time is equal to its genetic variance injtness at that time. ’ And also in these words (1930, p. 46; 1958, p. 50):
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George Price (1972) studied this question.
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