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The theory of quantitative genetics bridges the gap between observable statistical properties of a character and the genetic and environmental factors which are thought to determine its expression. This book covers a wide range of genetic topics from Mendelian genetics to selection theory in breeding. It looks at the decomposition of the genotypic value into components representing various types of gene action; estimation of the components of variance and other quantities generated by this decomposition, and stochastic effects of finite population size.
M. G. Bulmer (Sun,) studied this question.