Key points are not available for this paper at this time.
F. Stuart Chapin III, Douglas A. Johnson, Jay D. McKendrick, Seasonal Movement of Nutrients in Plants of Differing Growth Form in an Alaskan Tundra Ecosystem: Implications for Herbivory, Journal of Ecology, Vol. 68, No. 1 (Mar., 1980), pp. 189-209
Chapin et al. (Sat,) studied this question.