F. Stuart Chapin III, Douglas A. Johnson, Jay D. McKendrick, Seasonal Movement of Nutrients in Plants of Differing Growth Form in an Alaskan Tundra Ecosystem: Implications for Herbivory, Journal of Ecology, Vol. 68, No. 1 (Mar., 1980), pp. 189-209
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Chapin et al. (1980) studied this question.
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