Anaspis Geoffroy, 1762 subgenus Spanisa Emery, 1876 (Coleoptera: Scraptiidae) as Paleogene relicts

Anaspis (Spanisa) solodovnikovi sp. nov., the second known representative of its genus and subgenus from the late Eocene Rovno amber of Ukraine, is described and illustrated. The taxonomic position is confirmed, and supplementary description and images are provided for the Eocene Baltic amber A. (Silaria) parva Abdullah, 1964. A key to extant and extinct species of the subgenus Spanisa Emery, 1876 is presented for the first time. Morphology and new taxonomic position of Anaspis (incertae sedis) subhumerosa Franciscolo, 1954 are briefly discussed; this species is excluded from the subgenus Spanisa. Extant Spanisa species are allopatric, but in Rovno amber they are sympatric likely because of their adaptation to the ‘extinct’ equable Eocene climate. New faunistic records are provided for the extant A. (Spanisa) labiata A. Costa, 1854 and A. (Spanisa) subtilis Hampe, 1871.